Gard路易艾玛

83AnupdatedchecklistoftheextantWesternPalaearcticDryininae(Hymenoptera,Dryinidae)AdalgisaGuglielmino1,MassimoOlmi2,Jing-xianLiu3,MarioContarini11DepartmentofAgricultureandForestSciences(DAFNE),UniversityofTuscia,Viterbo,Italy2TropicalEntomologyResearchCenter,Viterbo,Italy3DepartmentofEntomology,SouthChinaAgriculturalUniversity,Guangzhou,ChinaCorrespondingauthor:Jing-xianLiu(liujingxian@yahoo.
com;liujingxian@scau.
edu.
cn)Academiceditor:MichaelOhl|Received21January2019|Accepted23July2019|Published5September2019http://zoobank.
org/7324B70E-EE52-4825-9DC8-51347B30DBF0Citation:GuglielminoA,OlmiM,LiuJ-x,ContariniM(2019)AnupdatedchecklistoftheextantWesternPalaearcticDryininae(Hymenoptera,Dryinidae).
ZooKeys874:83–99.
https://doi.
org/10.
3897/zookeys.
874.
33226AbstractAchecklistof20extantspeciesofDryininae(Hymenoptera,Dryinidae)fromtheWesternPalaearcticsubregionispresented.
KeywordsChecklist,distribution,ChrysidoideaIntroductionPincerwasps(Hymenoptera,Dryinidae)areparasitoidsandoftenalsopredatorsofAuchenorrhyncha(Hemiptera)(Olmi1984).
Thefamilyincludes50generaand16subfamilies(OlmiandXu2015;Tribull2015).
InthePalaearcticregion,thesubfamilyDryininaeisrepresentedbytwoextantgenera,DryinusLatreille,1804andPseudodryinusOlmi,1991.
PseudodryinusisknownonlyfromtheEasternPalaearcticsubregionandDryinusfrombothPalaearcticsubregions,EasternandWestern(OlmiandXu2015).
AreviewoftheWesternPalaearcticDryininae(Hymenoptera,Dryinidae)waspublishedbyOlmi(1984),andhelistedatotalofninespecies.
However,inthelast25yearsmanyadditionalpapersontheWesternPalaearcticfaunahavebeenpublished,sothatthenumberofspecieshasincreasedto20,andtheneedtodevelopanewchecklistofWesternPalaearcticDryininaebecameevident.
TheobjectiveofthischecklististoeasefurtherstudiesonPalaearcticdryinids.
ZooKeys874:83–99(2019)doi:10.
3897/zookeys.
874.
33226http://zookeys.
pensoft.
netCopyrightAdalgisaGuglielminoetal.
ThisisanopenaccessarticledistributedunderthetermsoftheCreativeCommonsAttributionLicense(CCBY4.
0),whichpermitsunrestricteduse,distribution,andreproductioninanymedium,providedtheoriginalauthorandsourcearecredited.
CHECKLISTLaunchedtoacceleratebiodiversityresearchApeer-reviewedopen-accessjournal84MaterialandmethodsThepresentpapertreatsallextantDryininae(fossilspeciesareexcluded)presentintheWesternPalaearcticsubregion,i.
e.
,accordingtoVignaTagliantietal.
(1992,1999),thepartofthePalaearcticregionsituatedinEuropeandAsiawesttotheUralMountainsandCaspianSea,fromtheAzoresandCanaryIslandstoIran(included).
Thebordersarenotalwaysobviousandnatural.
TheeasternboundaryrunsalongtheUralMoun-tainsandtheeasternbankofCaspianSea,reachingIran.
TheRussianregionsituatedimmediatelyeastofUralMountainsinpartsofKazakhstan,Turkmenistan,andIranshouldbeconsideredtransitioncountrytotheEasternPalaearcticsubregion,whereasalargepartoftheArabianPeninsulashouldbeconsideredatransitionareatotheAfro-tropicalregion.
Allthesetransitionareasareconsideredinthischecklist.
TheknowledgeofthedryinidslivingintheWesternPalaearcticsubregionisbroadlyinsufficient,sothatthischecklistwillneedtobeupdatedinthefuturefollowingfurtherresearch.
DistributionaldataofDryininaeintheWesternPalaearcticregionwerecompiledanalysingalltheavailablepublications,inadditiontomanyunpublishedrecordsob-tainedbyidentifyingmaterialbelongingtovariousinstitutions.
Allthelocalitiescitedinthischecklist,exceptthatfromBelaruscitedbySh-lyakhtyenok(2013)(seeDryinuscollaris(Linnaeus)),werecheckedbytheauthorsbyexaminingpersonallyallthespecimens.
Theexaminedspecimensaredepositedinthefollowingcollections:AECChristophSaure'scollection,Berlin,Germany.
AMNHAmericanMuseumofNaturalHistory,NewYork,USA.
ASMAlexanderShlyakhtenok'scollection,Minsk,Belarus.
BNCBenotNusillard'scollection,MontbouchersurJabron,France.
BWCBogdanWiniowski'scollection,OjcówNationalPark,Ojców,Poland.
CASCaliforniaAcademyofSciences,SanFrancisco,California,USA.
CIRADCentreInternationaldeRechercheAgricolepourleDéveloppement,Montpellier,France.
CNCCanadianNationalCollectionofInsects(CNCI),Ottawa,Canada.
DEISenckenbergDeutschesEntomologischesInstitut,Müncheberg,Germany.
DEUWDepartmentofEntomology,UniversityofWageningen,theNetherlands.
DISAFADipartimentodiScienzeagrarie,forestaliealimentari,UniversityofTo-rino,Grugliasco,Torino,Italy.
DPPZDepartmentofPlantProtection,CollegeofAgriculture,UniversityofZabol,Iran.
ENSAMcoleNationalSupérieureAgronomique,Montpellier,France.
FBWForstlicheVersuchs-undForschungsanstaltBaden-Württemberg,Freiburg,Germany.
FSAEFacultédesSciencesAgronomiquesdel'tat,Gembloux,Belgium.
GLPCGianlucaParise'scollection,Carignano,Torino,Italy.
GNCGranNilsson'scollection,c/oDepartmentofZoophysiology,UppsalaUniversity,Uppsala,Sweden.
85GPCGuidoPagliano'scollection,Torino,Italy.
HMOHopeMuseum,Oxford,England,UnitedKingdom.
HTSHubertTussac'scollection,Cahors,Lot,France(nowc/oMuseumd'Histoirenaturelle,Genève,Switzerland).
IGCIliaGjonov'scollection,Sofia,Bulgaria.
IRSNInstitutRoyaldeSciencesNaturellesdeBelgique,Bruxelles,Belgium.
JBZCJavierBlasco-Zumeta'scollection,PinadeEbro,Zaragoza,Spain.
JTBCJohnT.
Burn'scollection,Sacriston,England,UnitedKingdom.
LOHCLarsOveHansen'scollection,Drammen,Norway.
MBCManuelBaena'scollection,Cordoba,Spain.
MCNTNMuseodeCienciasNaturales,SantaCruz,Tenerife,CanaryIslands,Spain.
MCSNGMuseoCivicodiStoriaNaturale"GiacomoDoria"diGenova,Italy.
MCSNVMuseoCivicodiStoriaNaturale,Verona,Italy.
MHNGMuséumd'HistoireNaturelle,Genève,Switzerland.
MLUHWMartin-Luther-Universitt,Halle-Wittenberg,Germany.
MNCNMMuseoNacionaldeCienciasNaturales,Madrid,Spain.
MNHNMuséumNationald'HistoireNaturelle,Paris,France.
MMBMoravianMuseum,Brno,CzechRepublic.
MOLCMassimoOlmi'scollection,Viterbo,Italy.
MSCMassimilianoSpinola'scollection,c/oMuseoRegionalediScienzeNatu-rali,Torino,Italy.
MSCSMartinSchwarz'scollections,c/oInstitutfürZoologie,Salzburg,Austria.
MSNTCMuseodiStorianaturaleedelTerritorio,UniversitàdiPisa,Calci,Italy.
MZUNMuseodiZoologiadell'Università,Napoli,Italy.
NHMUKNaturalHistoryMuseum,London,UnitedKingdom.
NMNHNationalMuseumofNaturalHistory,Budapest,Hungary.
NMPCNationalMuseum(NaturalHistory),Praha,CzechRepublic.
NMWNaturhistorischenMuseum,Wien,Austria.
OLLObersterreichischesLandesmuseum,Linz,Austria.
PNLPierre-NicolasLibert'scollection,Somme-Leuze,Belgium.
RNHLRijksmuseumvanNatuurlijkeHistorie,Leiden,TheNetherlands.
SVCSimoVnnen'scollection,Vantaa,Finland.
SZCPierLuigiScaramozzino'scollection,Pisa,Italy.
USNMNationalMuseumofNaturalHistory,Washington,DC,USA.
VVCVeliVikberg'scollection,Turenki,Finland.
WHCPaulWhitehead'scollection,MoorLeys,England,UnitedKingdom.
YUICYeungnamUniversityInsectCollection,DepartmentofBiology,Yeung-namUniversity,Kyongsan,SouthKorea.
ZILZoologicalInstitute,Lund,Sweden.
ZMKZoologiskMuseum,Copenhagen,Denmark.
ZMMZoologicalMuseumofMoscowUniversity,Moscow,Russia.
ZMUHZoologicalMuseumoftheUniversity,Helsinki,Finland.
86ChecklistoftheextantWesternPalaearcticDryininaeHaliday,1833GenusDryinusLatreille,18041.
Dryinusalbrechti(Olmi)RichardsidryinusalbrechtiOlmi1984:909.
Dryinusalbrechti(Olmi):Olmi1999:192.
SPAIN:CanaryIslands:Fuerteventura,LasPeitas(MNCNM)(Olmi1984);Lanzarote,ElRiscodeFamara(AMNH)(Olmi1984);Tenerife,Orotava(ZMUH)(Olmi1984).
Distribution:Spain.
2.
DryinusbalearicusOlmiDryinusbalearicusOlmi1987:418;Olmi1999:209.
SPAIN:BalearicIslands:Ibiza,5kmNSanJosé(NHMUK,AMNH)(Olmi1987).
ContinentalSpain:HuescaProv.
,nearTorla,Fanlo(NHMUK).
TUNISIA:BarrageMellègue(MOLC).
Distribution:Spain,Tunisia.
3.
Dryinusberlandi(Bernard)ChelotheliusberlandiBernard1935:41;Olmi1984:609.
Dryinusberlandi(Bernard):Olmi1999:202.
FRANCE:Var,Fréjus,Saint-Raphalbeach(MNHN)(Bernard1935).
MOROC-CO:alongRoadP39,69kmMelilla,DarDriouch(MBC)(Olmi1999).
TUNISIA:10kmNofJendouba(OLL).
Distribution:France,Morocco,Tunisia.
4.
Dryinuscanariensis(Ceballos)ParadryinuscanariensisCeballos1927:101.
Dryinuscanariensis(Ceballos):Olmi1984:734;1999:184.
SPAIN:CanaryIslands:Gomera,SanSebastian,BarrancodeMarchar(MNCNM)(Olmi1984);Gomera,Chejelipes(NHMUK);Hierro,Frontera(MCNTN);LaPalma,MontaaBrena(OLL);Tenerife,BarrancoSantos(AMNH,MCNTN,MNCNM)(Olmi1984);Tenerife,DesembocaduradelBarrancodeTejina(AMNH);Tenerife,LaCuesta(MNCNM)(Ceballos1927);Tenerife,Tahodio(MNCNM)(Olmi1984);87Tenerife,Médano,LosCalderones(MCNTN,MNCNM)(Olmi1984);Tenerife,Mé-dano(MNCNM);Tenerife,Arico,MontaaAtalaya(MNCNM);Tenerife,Bajamar(MNCNM);Tenerife,LasMercedes(MNCNM);Tenerife,CarreteradeSanAndres,Jagua(AMNH);Tenerife,RaguoNegro(OLL).
EGYPT:Sinai,SaintCatherinearea(MOLC).
GREECE:RhodesIsland,ridgeNofPsinthos(NHMUK).
Distribution:Egypt,Greece,Spain.
5.
Dryinuscollaris(Linnaeus)SphexcollarisLinnaeus1767:946.
DryinusformicariusLatreille1805:228(synonymizedbyFittonetal.
1978).
CampylonyxampuliciformisWestwood1835:52(synonymizedbyOlmi1984).
Lestodryinusformicarius(Latreille):Kieffer1914a:20.
LestodryinuscorsicaeKieffer1914a:21(synonymizedbyOlmi1984).
Dryinus(Lestodryinus)formicariusLatreille:Haupt1932:15.
Dryinuscollaris(Linnaeus):Olmi1999:185;OlmiandXu2015:133.
AUSTRIA:Niedersterreich,Piesting(NMW);Obersterreich,Hinteraigen,EAi-bach/Donau,48°24'N,13°57'E(OLL);Salzburg,Werfen(RNHL)(Olmi1984);Steiermark,O-Steiermark,EWeiz,Hoferberg(OLL);Wien,Dornbach(MNHN)(Olmi1984).
BELARUS:PolyeseRadiacyonno-ekologicheskyzapovednik,Dron-ki(ASM)(Shlyakhtyenok2013).
BELGIUM:Brabant,ForêtdeSoignes(IRSN);Liège,Flémalle-HauteauxRoches(IRSN);Namur,Ave-et-Auffe,Thérimont(IRSN);Namur,Somal(PNL).
CROATIA:Istra,Opatija(NMNH)(Olmi1984);Krapina(NMNH)(Olmi1984).
FRANCE:AlpesdeHaute-Provence,Digne(MNHN)(Olmi1984);Corse(ENSAM)(Olmi1999);Gard,Bez-et-Esparon(CIRAD)(TussacandOlmi1998);Haute-Garonne,Clermont-le-Fort(HTS)(TussacandOlmi1998);Haute-Garonne,Toulouse,L'Isle-Jourdain(FSAE);Haute-Loire,LePuy(MNHN)(Olmi1984);Haute-Savoie,Bossy-Frangy(MHNG);Hérault,Saint-Gély-du-Fesc(MNHN)(Olmi1984);Hérault,Mons-la-Triviale,Gorgesd'Héric(CIRAD)(TussacandOlmi1998);Hérault,Saint-Guilhem-le-Désert(MNHN)(Olmi1984);Landes,Mont-de-Marsan(MNHN)(Olmi1984);Pyrénées-Orientales,ForêtdeBoucheville(DEUW);Rhne,Lyon(MNHN)(Westwood1835);Sane-et-Loire,LesGerraux(MNHN)(Olmi1984);Var,Hyères(MNHN)(Olmi1984);Var,Toulon(MNHN)(Olmi1984);Var,Sainte-Baume(MNHN);Vaucluse,nearBédoin(NHMUK).
GERMANY:Baden-Württemberg,Baden,Freiburgi.
B.
,BechtalerWald,48°12'N,07°42'E(FBW);Baden-Württemberg,FreiburgimBreisgau,Mooswald-Nord(AEC);Nordrhein–Westfalen,Aix-la-Chapelle(=Aachen)(KiefferandMarshall1905).
HUNGARY:Borsod-Abaúj-Zempléncounty,Cserépfalu,Hór-vlgy(NMNH,AMNH)(Szllsi-TóthandGyrgy2009).
ITALY:Campania,NapoliProv.
,Napo-li(KiefferandMarshall1905);EmiliaRomagna,BolognaProv.
,Gaibola(AMNH)(Olmi1984);Liguria,GenovaProv.
,S.
Olcese(NHMUK)(Olmi1984);Piemonte,CuneoProv.
,Valmala,alongCombadiValmala,PonteParasacco(IRSN);Piemonte,88TorinoProv.
,Rosta(AMNH)(Olmi1984);Piemonte,TorinoProv.
,Strambino(DIS-AFA)(Olmi1999);Puglia,TarantoProv.
,Mottola,S.
Basilio(Móczár1965);Sicilia,CataniaProv.
,Bronte,Maletto,Mt.
Etna,ContradaPaviglione(AMNH)(Olmi1999);Toscana,LuccaProv.
,Lucca(KiefferandMarshall1905);Toscana,LuccaProv.
,LidodiCamaiore(MOLC)(Olmi2005b);TrentinoAltoAdige,BolzanoProv.
,Bolzano(Schmiedeknecht1907).
MONTENEGRO:HercegNovi(=CastelnuovodiCat-taro)(DEI);Zelenika(NMNH)(Olmi1984).
POLAND:EastbankofOderRiver,10kmNofCedynia,Bielinek(=Bellinchen)(Haupt1932,asDryinus(Lestodryi-nus)formicarius).
SLOVAKIA:SWSlovakia,LittleCarpathians(MaléKarpaty),nearconfluenceDanubeandMoravaRivers,DevínskaKobylaHill(Lukás1998).
SPAIN:BalearicIslands:Mallorca,PortoCristo(RNHL).
ContinentalSpain:AlicanteProv.
,SierradeAltana(RNHL).
SWITZERLAND:Genève,Peney(MHNG)(Olmi1984);Genève(MSC)(Olmi1984);Genève,BoisdeCollex(MHNG);Genève,PlacedesNations(NMNH);Valais,Chteauneuf(MHNG);Ticino,Gandria(NMNH,AMNH)(Olmi1999).
THENETHERLANDS:Lexmond(RNHL);Neercanne,Cannerbos(RNHL)(DeRond2004).
TURKMENISTAN(Ponomarenko1978).
UNITEDKINGDOM:England:Berkshire,HighStandinghillWood,WindsorForest(NHMUK);Middlesex,Ruislip,VictoriaRoad(NHMUK);Surrey,BansteadDowns(NHMUK);Surrey,Shere(Capron1885);Surrey,Reigate,30ParkLaneEast(onlyphotographed,notcollected);WestKent,Cobham(Richards1939);WestKent,Eltham(JTBC);Worcestershire,MalvernHills(WHC)(Whitehead2010).
Distribution:Austria,Belarus,Belgium,Croatia,France,Germany,Hungary,Italy,Montenegro,Poland,Slovakia,Spain,Switzerland,theNetherlands,UnitedKingdom,inadditiontoTurkmenistan(transitioncountrytoEasternPalaearcticsubregion).
6.
DryinuscorsicusMarshallDryinuscorsicusMarshall1874:207;OlmiandXu2015:133.
Mesodryinuscorsicus(Marshall):Kieffer1907:10.
MesodryinusescorialensisCeballos1927:102(synonymizedbyOlmi1984).
Richardsidryinuscorsicus(Marshall):Móczár1965:377.
CYPRUS:Limassol(NHMUK)(Olmi1984).
FRANCE:Alpes-Maritimes,Breil-sur-Roya,ColdesTermes(CIRAD);Aude,Brouilla(BNC);Bouches-du-Rhne,Aix-en-Provence(NHMUK)(Olmi1984);Corse,Ajaccio,Campoloro(NMNH)(Olmi1984);Drme,Montségur-sur-Lauzon(HTS);Drme,Mévouillon(BNC);Drme,S.
teJalle(RNHL);Drme,Séderon,Coldel'Hommemort(AMNH);Drme,ColdeMacuègne(NHMUK);Gironde,Barsac(HTS)(TussacandOlmi1998);Hérault,Cazevieille(MNHN)(TussacandOlmi1998);Hérault,Montpellier(HTS)(TussacandOlmi1998);Hérault,Grabels(HTS);Hérault,LaFigarède(MNHN)(Olmi1984);Hérault,St.
Gély-du-Fesc(MNHN)(Olmi1984);Hérault,BalliarguetCSIRO,43°41.
12'N,03°62.
24'E(CNC);Haute-Garonne,Castelmaurou(HTS)(TussacandOlmi1998);Lot,Cahors(HTS)(TussacandOlmi1998);Lot,LeMontat(HTS);Var,nearSt.
Zach-89arie(NHMUK);Vaucluse,Sérignan(MNHN)(Olmi1984);Vaucluse,Lagarded'Apt,MtStPierre(BNC).
GREECE:Olympia,Ilia(NHMUK)(Olmi1984);Peloponìsos,Monemvasia(ZMK).
HUNGARY:Somogycounty,Kaposvár(NMNH)(Olmi1984).
ITALY:Calabria,CrotoneProv.
,Sila,alongroadfromPagliarelletoMt.
Gariglione,about9.
7kmfromPagliarelle,39°07.
382'N,16°41.
553'E(MOLC);FriuliVeneziaGiulia,TriesteProv.
,VillaOpicina(DEI)(Olmi1984);EmiliaRomagna,ForlìProv.
CampignaForest(MCSNV)(Olmi1999);Toscana,PisaProv.
,Lajatico,43°27.
86'N,10°40.
73'E(MOLC)(Olmi2005b).
KAZAKHSTAN:Tchimkentobl.
,KaratauRidgenearSuzak(ZMM)(PonomarenkoandOlmi2006).
SPAIN:Barcelona,Palamos(SZC)(Olmi1984);Murcia,nearManzarron(NHMUK)(Olmi1984);Murcia,SierradeEspua,nearTotana(NHMUK)(Olmi1984);Madrid,ElEscorial(MNCNM)(Ce-ballos1927);Granada,Cubillas(AMNH,NHMUK)(Olmi1984);Granada,Nerja(NHMUK);Castellon,Benicasim(NHMUK);10kmfromAbejar,Soria(RNHL);Ali-cante,Jávea(HTS);Zaragoza,PinadeEbro,Monegros(HTS,JBZC)(Olmietal.
1998).
Distribution:Cyprus,France,Greece,Hungary,Italy,Spain,inadditiontoKa-zakhstan(transitioncountrytoEasternPalaearcticsubregion).
7.
Dryinusdayi(Olmi)MesodryinusdayiOlmi1984:1003.
Dryinusdayi(Olmi):Olmi1999:204.
GREECE:Thessalia,Kalambaka(NHMUK)(Olmi1984).
Distribution:Greece.
8.
DryinusdelvareiOlmiDryinusdelvareiOlmi1998:72.
ALBANIA:MirditDistrict,Salit(MOLC).
ITALY:Toscana,ArezzoProvince,Upac-chi,43°30'N,11°59'E(MSCS);Toscana,GrossetoProv.
,MaremmaNaturalPark,42°38.
44'N,11°04.
42'E(MSNTC)(Olmi2005b).
TURKEY:18kmNWKorkuteli(AMNH)(Olmi1998).
Distribution.
Albania,Italy,Turkey.
9.
DryinusgharaeiiOlmiDryinusgharaeiiOlmi2005a:207;OlmiandXu2015:144.
IRAN:IlamProvince,ChogasabzRegion,Ilam(MOLC)(Olmi2005a).
Distribution.
Iran.
9010.
Dryinusgryps(Reinhard)ChelotheliusgrypsReinhard1863:410.
Dryinusgryps(Reinhard):DallaTorre1898:544;Olmi1995:5.
FRANCE:Bouches-du-Rhne,Fonscolombe(NHMUK);Drme,Montségur-sur-Lauzon(BNC);Gard,Ussel-Goudargues(AMNH)(Olmi1984);Hérault,Montag-nac,MasdeLinares(HTS)(TussacandOlmi1998);Lot,Cahors(HTS);SouthernFrance(MNHN)(Olmi1984).
ITALY:Sicilia,SiracusaProvince,Lentini(AMNH)(Olmi1999);Toscana,LivornoProvince,nearPiombino,Salivoli,42°56.
79'N,10°30.
20'E(MOLC)(Olmi2005b);Toscana,PisaProvince,MonteverdiMarittimo,43°09.
59'N,10°43.
24'E(MOLC)(Olmi2005b);Trentino-AltoAdige,Bolzano(Rein-hard1863).
SPAIN:Zaragoza,PinadeEbro,LosMonegros(HTS)(Olmietal.
1998);Madrid,ElPardo(MNCNM);Catalua,Tarragona,ElPerello(HTS).
TURKEY:Konya,Meram(OLL).
Distribution.
France,Italy,Spain,Turkey.
11.
Dryinusibericus(Olmi)AlphadryinusibericusOlmi1990:137.
Dryinusibericus(Olmi):Olmi1999:208.
SPAIN:Murcia,AlbaceteProv.
,nearMolinicos,ElPardal(MNHN)(Olmi1990);Granada,SierradeCazorla,Vacillo(HTS)(Olmi1999).
Distribution.
Spain.
12.
Dryinusmaroccanus(Olmi)RichardsidryinusmaroccanusOlmi1984:910.
Dryinusmaroccanus(Olmi):TussacandOlmi1998:488;Olmi1999:193.
ALGERIA:Oran(MHNG,MOLC)(Olmi1984).
FRANCE:Alpes-Maritimes,Val-bonne(BNC);Hérault,Cazevieille(CIRAD)(Olmi1999).
MOROCCO:Tangeri(MHNG)(Olmi1984).
SPAIN:Madrid,ElPardo,ElGoloso(AMNH)(Olmi1999).
Distribution.
Algeria,France,Morocco,Spain.
13.
DryinusnigerKiefferDryinusnigerKieffer1904:352;Olmi1999:206.
91Mesodryinusniger(Kieffer):KiefferandMarshall1906:497;Olmi1984:1005.
MesodryinusbrittanicusRichards1939:228(synonymizedbyRichards1953).
ALBANIA:Arras,10kmNWPeshkopi(OLL)(Olmi1994).
CYPRUS:Cherkes(NHMUK)(Olmi1984).
CZECHREPUBLIC:CentralBohemia,Celakovice,Lipo-vka(NMPC)(Macek2007);OrientalBohemia,ZeleznéhoryPLA,ZlatáloukaNa-tionalReserve(NMPC)(Macek2007).
DENMARK:SouthJutland,Sotrup(ZMK)(Olmi1994).
FINLAND:Satakunta,Eurajoki(VnnenandVikberg2007)(SVC,VVC).
FRANCE:Haute-Garonne,Castelmaurou(AMNH,MNHN)(TussacandOlmi1998);Lot,Lavercantière(HTS)(TussacandOlmi1998);Lot,Cahors(HTS);Vaucluse,MontVentoux,Malaucène(MNHN).
GERMANY:Rheinland-Pfalz,Gn-nersdorf(CllnandSorg2001).
GREECE:Peloponnesus,Monemvasia(ZMK).
ITA-LY:Campania,SalernoProv.
,VallodellaLucania(MCSNG)(Kieffer1904);Liguria,Genova(MCSNG)(Olmi1984);Piemonte,CuneoProv.
,Valdieri(AMNH)(Olmi1999);Piemonte,VercelliProv.
,Piode,AlpeMeggiana(MOLC).
NORWAY:InnerTelemark,Notodden,Lisleherad(LOHC)(HansenandOlmi1996;Olmi1994).
SLOVAKIA:SouthernSlovakia,Senec,Ciernavodariver(NMPC)(Macek2007).
SWEDEN:Smland(ZIL)(Olmi1994);Vrmland,Ekshrad(CNC)(Olmi1984);Vstmanland,Krrbo,Solbacken(GNC)(Olmi1994).
THENETHERLANDS:ZuidHolland,Lexmond(DeRond2004);Gelderland,Kesteren,Lienden(DeRond2004).
UNITEDKINGDOM:England:Dorset,GlanvillesWootton,f#holotypeofM.
brittanicus(HMO)(Richards1939);Northants,Ayno,21–25.
VI.
1945,R.
B.
Ben-sonleg.
,1f#(NHMUK)(Olmi1984);Oxfordshire,Otmoor,19.
VII.
1961,M.
W.
R.
deV.
Grahamleg.
,1f#(NHMUK).
Distribution.
Albania,Cyprus,CzechRepublic,Denmark,Finland,France,Ger-many,Greece,Italy,Norway,Slovakia,Sweden,TheNetherlands,UnitedKingdom.
14.
DryinussanderiOlmiDryinussanderiOlmi1984:731;Olmi1999:211.
Alphadryinussanderi(Olmi):Olmi1991:284.
BULGARIA:Melnik(AMNH)(Olmi1984);Sandanski(=Liljanovo)(MMB);UpperThracianPlain,Besaparihills,Novoselovill.
,42.
0974N,24.
4690E(IGC)(Lapeva-Gjonovaetal.
2018).
CYPRUS:10kmWCapeGréko,AyiaNapa(ZMK).
FRANCE:Alpes-Maritimes,Moulinet,SentierColdeTurini-Faysset,43°58.
51'N,07°24.
40'E(CIRAD);Drme,Séderon,Coldel'Hommemort(AMNH)(Olmi1999);Hérault,Grabels(HTS)(TussacandOlmi1998).
ITALY:Piemonte,TorinoProv.
,Susa,Gia-glione(AMNH)(Olmi1999).
RUSSIA:EuropeanRussia:OrenburgDistrict,Orsk(Ponomarenko1992:asRichardsidryinusalbrechtiOlmi).
Distribution.
Bulgaria,Cyprus,France,Italy,Russia.
9215.
DryinustamaricicolaRakhshani&OlmiDryinustamaricicolaRakhshaniandOlmiinDerafshanetal.
2016:412.
IRAN:SistanandBaluchestanProv.
,ZabolCounty,Zabol(MOLC)(Derafh-sanetal.
2016).
Distribution.
Iran.
16.
DryinustarraconensisMarshallDryinustarraconensisMarshall1868:204;Olmi1984:742.
DryinusszepligetiiKiefferinKiefferandMarshall1905:77(synonymizedbyOlmi1984).
Plastodryinusszepligetii(Kieffer):KiefferandMarshall1906:496.
Lestodryinustarraconensis(Marshall):Kieffer1914a:21.
LestodryinusgregoriHoffer1936:164(synonymizedbyMóczár1965).
LestodryinusbidensHaupt1937(synonymizedbyOlmi1984).
DryinusszepligetiiNecKieffer:Ponomarenko1981:879.
BULGARIA:Damianitsa,8kmSofSandanski(CAS);Sandanski(=Liljanovo)(MMB);SlnoevBrjag(OLL);Nessebar(AMNH);Mt.
Strandzha,Izgrevvillage,42°08.
41'N,27°48.
37'E(IGC).
CROATIA:Dalmatia,Novi(NMNH)(Olmi1984).
CZECHREPUBLIC:SouthMoravia,Pouzdranyi(Hoffer1936;Móczár1965);Moravia,Kobylí(OLL);Moravia,SofBrno,Bratcice,49°03'N16°31'E(OLL);Moravia,Havraniky,Znojmo,48°49'N,15°59'E(OLL).
FRANCE:AlpesdeHaute-Provence,Simiane-la-Rotonde(HTS);Aude,Sallesd'Aude(NMNH);Bouches-du-Rhne,Fonscolombe(NHMUK);Gironde,Barsac(HTS)(TussacandOlmi1998);Loiret,Orléans(MNHN)(Olmi1984);Lot,Cahors(HTS)(TussacandOlmi1998);Haute-Garonne,Castelmaurou(HTS)(TussacandOlmi1998);Hérault,Gra-bels(HTS);Hérault,Montpellier(HTS)(TussacandOlmi1998);Hérault,Se-lagouLake(NHMUK);Vaucluse,nearSt.
Didier,GrangeNeuve(NHMUK);Vau-cluse,LesConstants,nearBédoin(NHMUK).
GERMANY:Baden-Württemberg,Mühlacker-Mühlhausen(DeRond,pers.
comm.
;seeOlmiandDeRond2001).
GREECE:CorfuIsland,KatoKarakiana(JTBC)(Burn2011);RhodesIsland,Kre-mastiHills(NHMUK)(Olmi1984);RhodesIsland,Ixia(NHMUK);ChalkidikiPeninsula,AmolianiIsland(MCSNG).
HUNGARY:Veszprémcounty,Balatonken-ese(NMNH)(Olmi1984);Crkvenica(AMNH)(Olmi1984);Nógrádcounty,Ipol-ytarnóc(NMNH)(Szllsi-TóthandGyrgy2009);KiskunságNationalPark,Bugacpuszta(AMNH).
IRAN:KermanProv.
,BamCounty,Sangemes,28°56'33.
44"N,58°07'52.
36"E(DPPZ)(Derafshanetal.
2016);KermanshahProv.
,Kermanshah,Moghoye,28°57'24.
18"N,58°06'34.
90"E(DPPZ)(Derafshanetal.
2016).
IRAQ:93Baghdad(OLL,USNM)(Olmi1984).
ITALY:Abruzzi,PescaraProv.
,Mt.
Maiella(MZUN)(Olmi1984);Calabria,CrotoneProv.
,NofPetilia,nearroadtoPagli-arelle,39°07.
333'N,16°47.
036'E(MOLC);EmiliaRomagna,ParmaProv.
,Parma(MZUN)(Olmi1984);Lazio,ViterboProv.
,Capodimonte(AMNH)(Olmi1999);Lazio,ViterboProv.
,Sutri(MOLC);Liguria,LaSpeziaProv.
,Vernazza,44°08.
36'N,09°41.
73'E(MOLC);Liguria,SavonaProv.
,PietraLigure(GPC)(Olmi1984);Pie-monte,AlessandriaProv.
,Gavi(MCSNG)(Olmi1984);Piemonte,AlessandriaProv.
,MontaldodiCerrina(GLPC);Piemonte,CuneoProv.
,Valdieri,JuniperusphoeniceaReserve(AMNH)(Olmi1999);CuneoProv.
,S.
BenedettoBelbo(AMNH)(Olmi1984);TorinoProv.
,Carignano(Guglielminoetal.
2015);Puglia,LecceProv.
,S.
MariadiLeucaCape(MNHN)(Olmi1999);Puglia,TarantoProv.
,Castellaneta,BoscodeiTerzi,40°41'26,6"N,16°57'22,9"E(MOLC);Sardegna,SassariProv.
,Luras(MOLC)(Olmi2005b);Sicilia,CaltanissettaProv.
,S.
Cataldo(AMNH)(Olmi1984);Sicilia,CataniaProv.
,M.
Etna,Bronte,Maletto,ContradaPaviglione(AMNH)(Olmi1999);Sicilia,MessinaProv.
,Messina(ZMK)(Olmi1984);Toscana,GrossetoProv.
,Natu-ralParkofMaremma,42°38.
44'N,11°04.
42'E(MOLC)(Olmi2005b);Toscana,LivornoProv.
,Venturina(MOLC)(Olmi2005b);Toscana,LivornoProv.
,CapraiaIsland(MOLC)(Olmi2005b);Toscana,PisaProv.
,SanRossore,43°41'N10°39'E(MOLC)(Olmi2005b);Toscana,PisaProv.
,Cipollini(MOLC)(Olmi2005b);Tos-cana,PisaProv.
,MonteverdiMarittimo,43°09.
59'N,10°43.
24'E(MOLC)(Olmi2005b);Toscana,PisaProv.
,Lajatico,43°27.
86'N,10°40.
73'E(MOLC)(Olmi2005b);Toscana,PistoiaProv.
,MontecatiniTerme(AMNH)(Olmi1984);Umbria,PerugiaProv.
,Perugia(MCSNG)(Olmi1984);Valled'Aosta,AostaProv.
,Sarre(NHMUK)(Olmi1984).
MONTENEGRO:Sutomore(NMNH)(Olmi1984);nearKotor,Krasici(NHMUK).
MOROCCO:HighAtlas,25kmNofTaroudant,SebtTafraoute(NHMUK)(newrecord).
POLAND:EastbankofOderRiver,10kmNofCedynia,Bielinek(=Bellinchen)(MLUHW,MNHN)(Haupt1937);Wyzy-na,Malopolska,Rzezusniak/Golczy(BWC).
ROMANIA:Transilvania,Nagyenyed(NMNH).
RUSSIA:EuropeanRussia:VolgogradDistrict(Ponomarenko1978);FarEast:PrimorskijKraj,Evseyevka,15kmSEofSpassk(Ponomarenko1992).
SLO-VAKIA:SouthSlovakia,Stiavnické,HronskyBenadik(MMB).
SOUTHKOREA:GB,Gyeongsan-si,Dae-dong,Yeungnam-Univ.
,35°58'N128°47'E(YUIC).
SPAIN:Huesca,Torla(NMNH)(Marshall1868);Madrid,ElEscorial(NHMUK)(Olmi1984);Madrid,ElPardo,ElGoloso(MNCNM);Zaragosa,PinadeEbro,Monegros(HTS)(Olmietal.
1998);Castellon,Benicasim(NHMUK)(Olmi1984);Tarragona,Salou(AMNH,NHMUK)(Olmi1984);Granada,SierraNevada(AMNH,CNC);Alicante,Calpe(HTS);Almeria,Carboneras(RNHL);Soria,Ucero(RNHL).
TA-JIKISTAN:Kulyabobl.
,20kmENEPyandzh(ZMM)(PonomarenkoandOlmi2006).
TURKEY:Urfa,Halfeti(RNHL);Mugla,Kycegiz(RNHL);Hakkari,SWofYüksekova,Varegs,SatDag(RNHL);Pamphylia,WofAlanya(ZMK).
Distribution.
Bulgaria,Croatia,CzechRepublic,France,Germany,Greece,Hungary,Iran,Iraq,Italy,Montenegro,Morocco,Poland,Romania,Russia(incl.
94FarEast),Slovakia,Spain,Turkey,inadditiontoSouthKoreaandTajikistan(EasternPalaearcticsubregion).
17.
DryinustigaraeOlmiDryinustigaraeOlmi2008:365.
UNITEDARABEMIRATES:AbuDhabi,SweihanDistrict,AlAin(CNC)(Olmi2008).
Distribution.
UnitedArabEmirates.
18.
DryinusturcicusOlmiDryinusturcicusOlmi1991:259;1999:199.
TURKEY:HakkariDistrict,Hakkari(RNHL)(Olmi1991).
Distribution.
Turkey.
19.
DryinustussaciOlmiDryinustussaciOlmi1991:260.
FRANCE:Var,Vidauban(MNHN).
ITALY:Sardegna,SassariProv.
,Berchidda,40°47.
99'N,09°08.
87'E(MOLC)(Olmi2005b);Toscana,GrossetoProv.
,NaturalParkofMaremma,42°38.
17'N,11°04.
26'E(MOLC)(Olmi2005b);Toscana,PisaProv.
,MonteverdiMarittimo,43°09.
59'N,10°43.
24'E(MOLC)(Olmi2005b).
MO-ROCCO:about20kmNAgadir,Tarhazoute(MNHN)(Olmi1991).
SPAIN:Jaen,Alcaudete,SierradelAhillo(AMNH,MBC)(Olmi1999).
Distribution.
France,Italy,Morocco,Spain.
20.
DryinusyemenensisOlmiandVanHartenDryinusyemenensisOlmiandvanHarten2006:327.
OMAN:Dhofar,SalalahEast,Dahariz,17°01.
02'N,54°09.
32'E(MOLC).
YEMEN:AlLahima(MOLC)(OlmiandvanHarten2006);12kmNWofManakhah(MOLC)(OlmiandvanHarten2006);Al-Kowd(MOLC)(OlmiandvanHarten2006).
Distribution.
Oman,Yemen.
95DiscussionDryininaeoftheWesternPalaearcticsubregionareinsufficientlyknownfrommanypointsofview.
The20listedspeciesareknownmainlyonthebasisofonlyonesex(Derafshanetal.
2016;Olmi1999,2008;OlmiandvanHarten2006;OlmiandXu2015).
Infact,onlyfemalesareknownin11species(D.
berlandi,dayi,delvarei,gharaeii,gryps,ibericus,maroccanus,tigarae,turcicus,tussaci,yemenensis).
Bothoppositesexesareknowninsixspecies(D.
balearicus,collaris,corsicus,niger,sanderi,tarracon-ensis).
Inthreespecies,themalewasassociatedtothefemaletentatively,i.
e.
theasso-ciationisdoubtful(D.
albrechti,canariensis,tamaricicola).
Thissituationdependsonthelargemorphologicaldifferencesbetweenfemaleandmale,sothattheassociationoftheoppositesexesisimpossible,ifitisnotsupportedbyrearingsorDNAanalysis.
However,veryfewresearchersreardryinidsorstudytheirDNA.
Theknowledgeisinsufficientalsointheassociationofthespeciestotheirhosts.
DryinusspeciesareparasitoidsofFulgoromorpha(Guglielminoetal.
2013).
How-ever,intheWesternPalaearcticsubregion,thehostsareknownonlyinsixspecies(D.
balearicus,collaris,corsicus,niger,sanderi,tarraconensis).
Alsointhiscase,thesitu-ationdependsonthescarcityofrearings.
Fromthebiogeographicalpointofview,accordingtothecategoriespresentedbyVignaTagliantietal.
(1992,1999,thechorotypesofthe20specieslistedintheWesternPalaearcticsubregionarethefollowing:D.
albrechti(endemic,Macarone-sian);D.
balearicus(WesternMediterranean);D.
berlandi(WesternMediterranean);D.
canariensis(Mediterranean-Macaronesian);D.
collaris(Turan-European);D.
cor-sicus(Turan-European);D.
dayi(endemic,Greece);D.
delvarei(EasternMediterra-nean);D.
gharaeii(endemic,Iran);D.
gryps(SouthernEuropean);D.
ibericus(endem-ic,Spain);D.
maroccanus(WesternMediterranean);D.
niger(European);D.
sanderi(Turan-European);D.
tamaricicola(endemic,Iran);D.
tarraconensis(Asian-European);D.
tigarae(endemic,UnitedArabEmirates);D.
turcicus(endemic,Turkey);D.
tussaci(WesternMediterranean);D.
yemenensis(endemic,Yemen,Oman).
Eightspeciesoftheabovelistareconsideredendemicprovisionally,becausedryinidsareunderstudied,sotheirgeographicdistributioncouldbelarger.
OlmiandXu(2015)listed10speciesofDryinusandonespeciesofPseudodryinusfromtheEasternPalaearcticsubregion.
Sothedryinidpopulationofthetwosubre-gionshasaboutthesamenumericalstrength.
However,thecompositionisdifferent.
FewspeciesarepresentalsointheWesternPalaearcticsubregion,i.
e.
D.
collaris,D.
cor-sicus,andD.
tarraconensis.
AcknowledgementsManythankstoallcolleaguessendingonloanthespecimensstudiedinthepresentpaper.
ThisresearchwascarriedoutintheframeoftheMIUR(ItalianMinistryfor96Education,UniversityandResearch)initiative"Departmentofexcellence"(Law232/2016).
ThispaperwassupportedbytheNationalNaturalScienceFoundationofChina(No.
3147207;responsibleJingxianLiu).
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